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n An instrument with steel pins used to comb out flax or hemp. (defdate|from 15th c.)

n (usually|now|_|in the plural) One of the long, narrow feathers on the neck of birds, most noticeable on the cock. (defdate|from 15th c.)

n (fishing) A feather used to make a fishing lure or a fishing lure incorporating a feather. (defdate|from 17th c.)

n (usually|now|_|in the plural) By extension (because the hackles of a cock are lifted when it's angry), the hair on the nape of the neck in dogs and other animals; also used figuratively for humans. (defdate|from 19th c.)

n A plate with rows of pointed needles used to blend or straighten hair. (defdate|from 20th c.)

n A feather plume on some soldier's uniforms, especially the hat or helmet.

v To dress (flax or hemp) with a hackle; to prepare fibres of flax or hemp for spinning. (defdate|from 17th c.)

v (transitive) To separate, as the coarse part of flax or hemp from the fine, by drawing it through the teeth of a hackle or hatchel.

v (archaic|transitive) To tear asunder; to break into pieces.